My Work #
Studies over the past two decades seem to indicate that ocean currents at small scales (of order of magnitudes between 1km and 10km) have a strong influence on phytoplankton. However, precisely quantifying their impact on the ecosystem as a whole – that is, on regions typically spanning thousands of kilometers – is challenging.
PhD #
During my thesis, I attempted to quantify the impact of fronts (key structures at small scales) on phytoplankton in a large area around the Gulf Stream, using satellite imagery.

Gulf Stream meanders from space: Chlorophyll-a (top) and SST (bottom). MODIS-Aqua L2 data, 2007-04-22.
We detected fronts by computing an Heterogeneity Index from satellite sea surface temperature (Haëck et al. 20231, Liu & Levine 20162).
We then quantified the chlorophyll (also from satellite data) inside and outside these fronts. This work was made challenging by the presence of regions with different biogeochemical regimes and large-scale gradients within these regions. We show the importance of distinguishing the impact of fronts measured locally, and regionally by taking the impacted surface into account. We observe that the impact of fronts depends on their intensity. Lastly we were able to measure an earlier bloom onset with the fronts, by one to two weeks.

Local Chl-a excess over fronts and regional surplus taking the front surface into account Haëck et al. 2023, figure 9.
These findings are presented in Haëck et al. 20231 as well as in my PhD manuscript (in french).
Current #
I am continuing to work on refining the methodology that I established during my PhD. We have since applied it to data of phytoplankton composition generated by Roy El Hourany3. We found a population shift towards more diatoms and less prokaryotes at fronts, while the change in proportion of the other eukaryotic groups varies4. This shift accounts for up to a half of the large-scale community dissimilarity, underscoring the substantial influence of fronts on phytoplankton community composition.
Phytoplankton community composition for each biome over non-front, weak fronts and strong front conditions. Computed over 2002−2020. Lévy et al. 2025, figure 5.
It has also been applied in the California Current region5, with the aim (amongst other) of extending the results above to other areas. As part of the 4DMED-Sea project, this methodology is being applied to the mediterranean sea. Several avenues of study permitted by the novel 4DMED data are being considered: effect of fronts at depth, comparison of thermal, dynamical, and density fronts,…
The codes used for these studies are publicly available (gitlab.in2p3.fr/clementhaeck/submeso-color), and I am currently working to make them more accessible and easily reusable, thanks to two auxiliary projects: heterogeneity-index and data-assistant. Their application at a global scale is in development at gitlab.in2p3.fr/biofronts/submeso-color
I have also started compiling front-detection methods in a single Python package: fronts-toolbox.
Haëck C., Lévy M., Mangolte I., Bopp L. “Satellite Data Reveal Earlier and Stronger Phytoplankton Blooms over Fronts in the Gulf Stream Region”. Biogeosciences 20.9 (may 2023). doi:10.5194/bg-20-1741-2023 ↩︎ ↩︎
Liu X., Levine N. M. “Enhancement of Phytoplankton Chlorophyll by Submesoscale Frontal Dynamics in the North Pacific Subtropical Gyre”. Geophys. Res. Lett. 43.4 (2016). doi:10.1002/2015gl066996 ↩︎
El Hourany R., Abboud‐Abi Saab M., Faour G., Aumont O., Crépon M., Thiria S. “Estimation of Secondary Phytoplankton Pigments from Satellite Observations Using Self-Organizing Maps (SOMs)”. J. Geophys. Res. Oceans 124.2 (feb. 2019), p. 1357-1378. doi:10.1029/2018jc014450 ↩︎
Lévy M., Haëck C., Mangolte I., Cassianides A., El Hourany R. “Shift in phytoplankton community composition over fronts” Commun. Earth Environ. 6.1 (jul. 2025), p. 591. doi:10.1038/s43247-025-02553-1 ↩︎
Mangolte I., Lévy M., Haëck C., Ohman M.D. “Sub-frontal niches of plankton communities driven by transport and trophic interactions at ocean fronts”. Biogeosciences 20.15, (aug. 2023). doi:10.5194/bg-20-3273-2023 ↩︎